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However, many favoured competing explanations and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution.

Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh; instead, he helped to investigate marine invertebrates.

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The boy stood in awe of his overbearing father, whose astute medical observations taught him much about human psychology.One disgruntled singleton wrote: ‘You are everything that’s wrong in this world.